Betty has a hard time telling Daniel important news about her future; Amanda copes with the death of her dog; Marc wonders if he will ever have a successful relationship.

As Hilda’s wedding day approaches, Betty asks Daniel to be her plus-one, Ignacio invites Elena, and Amanda is set on fixing up Marc with her new client, soap opera star Spencer Cannon (Bryan Batt). Meanwhile, Justin weighs whether his own plus-one could

Betty brings Hilda on an assignment to cover London Fashion Week; Willie joins AA to get closer to Tyler and to double-cross Claire and Daniel; Bobby wonders why Justin is spending time with Austin.

After being knocked unconscious at a photo shoot, Betty has a dream in which her orthodontist (Kathy Najimy) shows her how her life would have been had she never worn braces.

Justin finds himself in an unexpected love triangle; Betty falls for a playwright she is profiling for Mode; Wilhelmina has an epiphany.

Betty is pressured into dating an annoying firefighter (Rich Sommer); Amanda persuades Claire to hire Tyler; Hilda meets Bobby's parents.

Daniel helps Betty to cover a low-priority show. Betty discovers a new designer, Marisa, and decides to promote her however she faces stiff competition from Marc. Amanda struggles to persuade Marc or Betty to vouch for her to Wilhemina about her clothing line. Tyler tries to find Claire in New York.

Wilhemina is livid when she lays eyes on drag queen impersonator - Wilhediva Hater. Betty and Daniel go through a rough patch as she wins an award for her blog. Daniel has the press hot on his heels. While Marc is pleased with himself when a famous director does a photo shoot with him.

Betty plans a karaoke party in her apartment building, however her building gets robbed. Marc loses something valuable. Meanwhile, Amanda tricks Daniel and Wilhelmina into attending a couples counseling session with Dr. Sparks.

Betty recalls everything that has made up her worst week in history, and also has job issues caused by taking up a new hobby. Hilda is treated to a week of luxury by Bobby, and Daniel is becoming suspicious of Mark's actions.

Betty wants to spend some time apart from Matt, so finds him something else to keep him occupied. Meanwhile, Hilda has a tough decision to make regarding her baby, and Mark's fling looks to be heading towards something more serious.

Ignacio prepares a romantic dinner for his Jean. Betty and Hilda take the Be-Shure instant pregnancy tests. Meanwhile, Hartley gets Denise Ludwig to fill in for Wilhemina. However Marc plans to bring Willie back. Claire with Amanda track down Tyler, the son she gave up for adoption.

Betty, Matt and Amanda's love triangle in the Bahamas reaches boiling point. Betty has trouble with a photo shoot when Penelope Graybridge steals the shoot from her enemy Wilhemina. Marc falls for super sexy Troy.

Daniel hands over the power to the Community of the Phoenix to run Mode. Betty steps in to help Daniel with Bennet. She comes up with an cunning plan with the help of Claire. Wilhemina goes out in public wearing sweats creating mayhem with the media trying to get a scoop.

Hilda has doubts over her relationship with Archie after seeing an old flame Bobby Talercio. Marc helps Justin to fit in high school by telling him to become friends with some mean girls. Elsewhere Daniel is lured into a trap by Bennett. Wilhemina tries to come up with money to help Nico out of trouble.

At Wilhemina's fund-raiser for Tibetan orphans Matt asks someone to be his partner. Daniel hires Natalie as his assistant, making Betty feel unneeded. Daniel finds out that Claire has a secret.

Betty has Marc as be her temporary assistant thanks to Wilhemina. Betty ends up covering the worst jobs in New York as an assignment but she needs to dress up as a hot dog and perform a Bollywood style dance. Marc has insecurities about how his idol at Vogue magazine will percieve him. Daniel is targeted by the leader of the Community.

Betty tries to get a scoop with Gucci's new designer, Evan York, through Hilda's customer, Sammy. Marc tries to stop her by getting Amanda and Matt to stage a showdown at Ignacio's restaurant. Daniel goes to a bereavement group and meets Natalie who shows him an interesting night. Wilhelmina flies to Bermuda to try and find Connor.

Betty begins to realise that her new job may not be all it's cracked up to be as she is working for her ex, is sharing an office with someone who doesn't like her, and Marc is trying to unseat her because he wants the job. Elsewhere, Daniel is lost without Betty at his side and he's also dealing with losing his wife. Wilhemina has got problems of her own - in and out of the office. Betty welcomes the chance to get to take charge of her first photo shoot, but when Daniel and Matt interfere, things don't go to plan. Amanda meets Daniel's new temp assisstant and who is very similar to herself, whilst Hilda needs help from Marc on how to deal with Justin.